Sir Henry Dale Research Fellow at Oxford University. His research focuses on investigating the patterns of gene flow in pathogen populations. He is particularly interested in phylogenetic methods of sequence analysis that combine genetic, spatial and ecological information. Specific questions involve (i) uncovering the spatiotemporal dynamics of human and animal pathogens, (ii) identifying factors underlying pathogen spread and dynamics at different scales (e.g. geographic regions, body compartments) and (iii) investigating the drivers of cross-species transmission and host shifts.